In APS, biological organisation is not merely spatial arrangement or structural complexity. It is organisation sustained through time.
Living systems persist by coordinating processes occurring at different temporal scales: metabolic turnover, repair, regulation, development, behaviour, reproduction, inheritance, ecological interaction, and evolution.
Temporal organisation therefore concerns the ordered relation among these processes. Some processes occur rapidly, others unfold slowly, but their biological significance depends on how they contribute to the continuing viability of the system.
This means that biological explanation cannot treat time as a neutral background. Time is part of the organisation being explained.
A biological system is not simply a thing that exists through time. It is a processual organisation whose persistence depends on the coordination, repair, transformation, and renewal of its own conditions of existence.